Suzume and Souta catch up to the capricious god Daijin after chasing him all over Japan, only to find the earthquake-inducing Worm he wast meant to seal away has broken free of its final binds. As certain disaster threatens to decimate Tokyo, Souta asks Suzume to help him make the ultimate sacrifice and take Daijin's place in the Ever-After. Now, desperate to save Souta from an eternity in the cold, Suzume sets off on one last trip back to the sight of her greatest tragedy in hopes of digging up memories that could provide the key to rescuing Souta-and to the door of her destiny.
